Match List-I (Evolutionists) with List-II (Achievements) and select the correct answer:

List-I	List-II
a. Haeckel	1. Emphasized the effects of different environments on isolated forms
b. Wagner	2. Suggested that an inner directive guides the course of evolution mainly independent of the environment
c. Guli	3. Pointed out the inevitability of divergence even without difference in environment
d. Karl Nageli	4. Constructed phylogeny of vertebrates on their embryological development

A. a-1, b-2, c-4, d-3

B. a-4, b-3, c-1, d-2

C. a-4, b-2, c-3, d-1

D. a-4, b-1, c-3, d-2

Answer: Option D