Sandal spike disease of sandal (Santalum album) is caused by

Sandal spike disease of sandal (Santalum album) is caused by MLO.
Spike disease caused by MLO (phytoplasma type) is the major disease of sandalwood. This disease is characterized by an extreme reduction in leaf size accompanied by stiffening and reduction of internode length.
Sandal spike causal agent is a pleomorphic microorganism and is the smallest organism capable of independent replication.